Understanding wolf ecology at landscape scales reveals how wolf welfare and ecosystem health are inseparably linked.
Wolf landscape welfare requires thinking across spatial scales that challenge human planning horizons. Individual wolf welfare — adequate territory, pack social integrity, prey access — requires landscape conditions that extend far beyond protected area boundaries. Lethal control that disrupts pack structure creates a welfare paradox: the social disruption from killing wolves may increase inexperienced predation attempts on livestock, worsening the conflict that motivated the killing. Coexistence approaches that maintain pack integrity while protecting livestock consistently outperform lethal control in both human-wildlife conflict reduction and wolf welfare outcomes.