Spotted hyenas live in complex matriarchal clans with sophisticated social hierarchies that shape individual welfare, access to resources, and survival outcomes.
Social rank profoundly affects hyena welfare - subordinate individuals have restricted access to food, experience more stress hormones, and have lower reproductive success. Clan conflicts can cause serious bite wounds that may become infected. Human persecution - retaliatory killing for livestock predation - causes injury, death, and disruption to clan social structure. Orphaned hyena cubs removed from their mother before 18 months have poor survival prospects due to the extended period of maternal care hyenas require.